Since 3.2.1 these characters could no longer precede an inline markup start-string or a bare URI or email address, although the reStructuredText spec allows them. So <https://example.com> was not a link and <jane-doe@example.com> became a link to doe@example.com. Also implement the rule that a start-string between an opening character and its closing counterpart, as in (*) or "*", is not inline markup.